van Herwaarden OA & van der Wal N 2002 Extinction time and age of an allele in a large finite population. Theor Popul Biol 61:311-318. c = (ln(8Nes) + (2 − 3h) ln(h) + (3h − 1) ln(1 − h) + γ) / (2sh(1 − h)) ... (19) for initial frequencies…