Uecker H & Hermisson J 2011 On the fixation process of a beneficial mutation in a variable environment. Genetics 188:915-930. Ne,t = Nt / (2ξ(t, Nt) + b(t, Nt) + d(t, Nt) + s(t, Nt)) ... (5) fixation probability dN(t) / dt = r(t, N(t))N(t)…